The 23-year-old came on for Paul Scholes after 70 minutes in a familiar holding position in midfield as England progressed to the quarter-finals with a thrilling 4-2 win against Croatia.
Of course, it's a role Ledley took up at Spurs for most of last season before moving back into defence for the last couple of games.
He was selected for England as a defender and was outstanding in the opener against France in John Terry's absence. Terry was back for the second game against Switzerland and remains first-choice centre-back alongside Sol Campbell.
Ledley King was back in action as England marched on in Euro 2004 last night - playing in midfield.